HKY85 (Hasegawa, Kishino & Yano, 1985) substitution model of nucleotide evolution.
Hasegawa M, Kishino H, Yano T (1985) Dating the human-ape splitting by a molecular clock of mitochondrial DNA. Journal of Molecular Evolution 22:160-174.
 
| type: beast.core.parameter.RealParameter |
| kappa parameter in HKY model |
| Required input |
 
| type: beast.evolution.substitutionmodel.Frequencies |
| substitution model equilibrium state frequencies |
| Required input |